Frank Buckles, Last Remaining American World War I Vet, Dies at 110

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U.S. President George W. Bush sits with World War I Veteran Corporal Fran Buckles March 6, 2008 (REUTERS / Jason Reed)

The last remaining World War I vet, Frank Buckles, died earlier Sunday morning at the old age of 110.

Part of the troop nicknamed “The Doughboys,” he served as a U.S. Army ambulance driver during the war.  He made history when he testified before Congress in 2009 over the construction of a monument on the National Mall, and has been profiled in photography projects documenting the last survivors of the “Great War.” While he never said much about his war experience, Buckles was drawn to the plight of the D.C. War Memorial for World War I veterans.

(More on TIME.com: See the making of the American G.I.)

Buckles died of natural causes; he had just turned 110 earlier this month. According to CNN and family, though, “he had slowed considerably since last fall.”

He will be buried in Arlington, Virginia following his wishes, alongside in the same cemetery where General John Pershing is buried. John Pershing was the commander in charge of American troops during World War I.

(More on TIME.com: See landscapes of World War I)